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What Is Microbial Diversity?
Facets of microbial diversity
What is diversity? Howexactly are organisms either similar to or different from each other? This seems like an easy question in the macroscopic world, but what about microbes?
Morphological diversity
Microbes are often divided by shape into rods, cocci, and spirals. Although these are the most common cell shapes, bacterial and archaeal cells also come in a wide range of other shapes: filaments (branched or unbranched), irregular, pleomorphic (different shapes under different conditions or even in the same culture), star-shaped, stalked, and many, many others.Haloquadratum is a flat, square organism, just like a bathroom tile (Fig. 1.1).
Individual cells of whatever shape can be found in a variety of multicellular arrangements, from simple pairs and tetrads to multicellular filaments, sheets, rosettes, and true multicellular organisms. Many species form highly structured multispecies mats that resemble the tissues of animals and plants that carry out complex biochemical transformations (Fig. 1.2).
Figure 1.1 The tile-shaped halophilic archaeonHaloquadratum walsbyi. (Source: Wikimedia Commons.) doi:10.1128/9781555818517.ch1.f1.1
Figure 1.2 Section of a stratified microbial mat from Guerrero Negro, Baja California. (Copyright 2007, American Society for Microbiology. Photo by John R. Spear and Norman R. Pace.) doi:10.1128/9781555818517.ch1.f1.2
Most bacteria and archaea measure 1 to 5 µm, but they range from 0.1 μm in thickness to over a millimeter. At the low end, it is hard to understand how everything that is needed for life could fit into the cell. At the high end, they can be easily seen without a microscope (Fig. 1.3).
Structural diversity
Many bacteria have “typical” gram-positive (single membrane, thick cell wall) or gram-negative (double membrane, thin cell wall) cell envelopes. However, there is wide variation even within these two major types. Many gram-positive bacteria have an outer membrane, made of mycolic acids rather than glycerol-phosphate esters. Many gram-negative bacteria lack the lipopolysaccharide layer. Many archaea and bacteria (both gram positive and gram negative) have an orderly protein coat, the S-layer (Fig. 1.4). In bacteria, cell walls are composed of peptidoglycan, but there is a surprising range of chemical variations within this type of material. Archaea do not have peptidoglycan cell walls, although some archaeal cell walls contain a related material, pseudomurein.
Figure 1.3 The bacteriumEpulopiscium fishelsoni (ca. 500 μm long) and four cells of the protistParamecium (ca. 100 μm long). (Courtesy of Esther Angert.) doi:10.1128/9781555818517.ch1.f1.3
Microbes have a wide range of external structures: flagella, pili, fibrils, holdfasts, stalks, buds, capsules, sheaths, and so on. They also have a wide variety of internal structures such as spores, daughter cells, thylakoids, mesosomes, and the nucleoid. In reality, microbial cells are just as structurally organized, and diverse, as are eukaryotic cells.
Figure 1.4 A negative-stain electron micrograph of the S-layer ofPyrobaculum aerophilum. Scale bar, 200 nm. (Courtesy of Reinhard Rachel.) doi:10.1128/9781555818517.ch1.f1.4
Metabolic diversity
Macroscopic eukaryotes are not metabolically diverse; they are either chemoheterotrophic (e.g., animals) or photoautotrophic (e.g., plants). Bacteria and archaea have a much